This game takes place after the events in Ape Escape 2001, where Spike had stopped Specter from taking over the world. After some time had passed, the Professor had left for a vacation leaving his granddaughter, Natalie, and Spike's cousin, Jimmy, to watch over the laboratory. The Professor has left them with one task: deliver a load of monkey pants to the monkeys in Monkey Park. Jimmy accidentally sends not only the monkey pants, but some Monkey Helmets as well, wrecking the laboratory in the process. Specter gets a hold of one, and yet again makes an army of monkeys bent on ruling the world. Under Natalie's orders, Jimmy aims to capture all the monkeys and stop Specter (due to the mess-up from earlier). However, Specter has supplied five monkeys, The Freaky Monkey Five, with Vita-Z Bananas, making them stronger and more intelligent than the average monkeys.
Jimmy manages to capture all of the Freaky Monkey Five, except Yellow Monkey. Before his fight with White Monkey, Pipotchi, a baby monkey accompanying Jimmy, has been kidnapped, in order to transfer the data in his Monkey Helmet (a re-engineered model which makes Pipotchi friendly) into a device called the Lethargy Laser. Rescuing Pipotchi, Jimmy makes his way through a monkey controlled military base, and confronts Specter, who is waiting for him with a redesigned version of his battle cruiser from the first game. Before the two can battle, Yellow Monkey, now a giant monster thanks to an overdose of Vita-Z bananas attacks the facility. Specter flees, and Jimmy manages to downsize and capture Yellow.
Jimmy, the Professor and Natalie then discover the purpose of the Lethargy Laser. By firing it at the globe, all humans worldwide will lose the will to fight back, making it easier for Specter and the monkeys to take over the world. Jimmy confronts Specter at his moonbase, who attacks him with a giant robot. Once defeated, Specter attempts to fire the laser, but Jimmy states that humans will never give up, and captures him. Jimmy and Pipotchi then flee the exploding moonbase.
Shortly afterwards, Specter escapes and goes into hiding. Jimmy then proceeds to capture all the remaining monkeys before the Professor discovers Specter's hiding place. Jimmy warps there, encountering Specter, and the two of them battle again. Specter loses and is once again captured. The game ends with Spike visiting the laboratory, which is again destroyed when Jimmy accidentally crashes a spaceship into it.
Like in the previous game, Ape Escape 2 features 3 minigames. But this time, instead of being unlocked only by collecting Specter Coins, the player has to pay for them with Gotcha Coins, collected throughout levels, via a vending machine called the Gotcha Box in the Travel Station. However, you still need to clear a specific level in order to make those available in the Gotcha Box.
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|Dance Monkey Dance||Clear Ninja Hideout|
|Monkey Soccer||Clear Enter The Monkey|
|Monkey Climber||Clear The Lost Valley|
Each of the game level's are connected through the Hub world, now the Travel Station. Jimmy now has to catch 300 monkeys.
|Liberty Island||Harley Q. Win (Secret Monkey)
|Castle Frightmare!||Arthur (Secret Monkey)
Arnie (Secret Monkey)
Leo (Secret Monkey)
Boxer (Secret Monkey)
Threedee (Secret Monkey)
Eddie the Falcon
Livingstone (Secret Monkey)
|The Blue Baboon||Helmut
Erin (Secret Monkey)
Mulder (Moldy NTSC)
Speedo (Speedy NTSC)
|Enter the Monkey||Jean Paul
Chan (Monkie Chan NTSC)
Bruce Monkee (Secret Monkey)
Miyazaki (Tai Shing NTSC)
John Ook (Monk NTSC)
Goku (Songokou NTSC) (Secret Monkey)
Daniel C. Ooke
Dolph (Secret Monkey)
Tommy (Secret Monkey)
Brucie (Spruce NTSC)
Woody (Woodward NTSC)
|Land of the Apes|
|The Lost World|
Play as Spike
If the player has successfully complete the game and catch all monkeys, Spike can be playable on a new file. Highlight the "New Game" option on the Main Menu, hold L1 and press Start to play as Spike. In this mode, all of the Monkeys become harder to catch. However, all of the Gadgets in the game become available from the beginning. There are also no cutscenes in this mode.
The music of the game was composed by Koji Hayama, managed by Don Makkou and Bungo Fujiwara of Two Five and data programmed by Masaaki Kaneko of Procyon Studio. The game's sound effects were created by Takashi Kanai and Junko Sano. The voice acting of the American version was recorded by Dan Rich and Jeremy Blaustein. The voice acting of the European version was recorded at Side UK on Great Titchfield Street in London, with Andy Emery serving as the voice director.
Ape Escape 2 was re-released as a PSN title in 2016, featuring trophies and upscaled graphics.
|Role||American Voice Actor||European Voice Actor||Japanese Voice Actor|
|Jimmy/Hikaru||Veronica Taylor||David Holt||Motoko Kumai|
|Pipotchi||Tara Sands||David Holt||Omi Minami|
|Natalie/Natsumi||Rachael Lillis||Rachel Preece||Jouji Yanami|
|Professor/Hakaze||Dan Green||Jonathan Keeble||Tomoko Kawakami|
|Specter||Gregory Abbey||Marc Silk||Chika Sakamoto|
|Blue Monkey/Ukki Blue||Gregory Abbey||Richard Pearce||Takahiro Sakurai|
|Yellow Monkey/Ukki Yellow||James Carter Cathcart||David Holt||Shoto Kashii|
|Pink Monkey/Ukki Pink||Veronica Taylor||Rachel Preece||Miyuki Matsushita|
|White Monkey/Ukki White||Gary Littman||Marc Silk||Kenichi Ogata|
|Red Monkey/Ukki Red||Gregory Abbey||Uncredited||Daisuke Gori|
|Spike/Kakeru||Dan Green||Richard Pearce||Fujiko Takimoto|
- The PlayStation 4 re-release of Ape Escape 2 has the European voice actors in the US. It is unknown why.
|Characters||Casi · Jake · Natalie · Professor · Specter · Spike|
|Gadgets||Dash Hoop · Magic Punch · Monkey Radar · R.C. Car · Sky Flyer · Slingback Shooter · Stun Club · Time Net · Water Net|
| || |
|Prehistoric Era/ The Lost Land||Fossil Field · Primordial Ooze · Molten Lava|
|Cenozoic Era/ Mysterious Age||Thick Jungle · Dark Ruins · Cryptic Relics|
|Dimension/Dimension X||Stadium Attack|
|Primitive Age/Oceana||Crabby beach · Coral Cave · Dexter's Island|
|Ice age/New Freezeland||Snowy Mammoth · Frosty Retreat · Hot Springs|
|Dimension/Dimension X||Gladiator Attack|
|Recent Past/Medieval Mayhem||Sushi Temple · Wabi Sabi Wall · Crumbling Castle|
|Present-Day/Futurama||City Park · Specter's Factory · TV Tower|
|Present-Day 2/Specter Land||Monkey Madness|
|Dimension/Dimension X||Peak Point Matrix|