The main villain of Ape Escape ~On Air~ Anime Season 1, He had wanted Specter's DNA for creating Biological weapons (which may explain why mutants appear, In Saru! Get You! Million Monkeys). He is the leader of the new species of Pipotrons created by a mad scientist who's greed over Haruka's father's disc. Pipotron J's species of primate is the Ring-tailed lemur. He also goes by the name Axel Dark-Key. Spoiler Alert: He gets revenge on Nazo, the mad scientist who created him, in Saru Getchu Million Monkeys, for killing his brothers. He gives him a taste of his own medicine by electro-shocking him. Whether he survived the attack is a mystery.

ApeEscapeTemplate ApeEscapeOTLTemplate
Characters Casi  · Jake  · Natalie  · Professor  · Specter  · Spike
Gadgets Dash Hoop  · Magic Punch  · Monkey Radar  · R.C. Car  · Sky Flyer  · Slingback Shooter  · Stun Club  · Time Net  · Water Net
Levels (/ Apes/ Enemies)
Hub World
Time Station
Prehistoric Era/ The Lost Land Fossil Field  · Primordial Ooze  · Molten Lava
Cenozoic Era/ Mysterious Age Thick Jungle  · Dark Ruins  · Cryptic Relics
Dimension/Dimension X Stadium Attack
Primitive Age/Oceana Crabby beach  · Coral Cave  · Dexter's Island
Ice age/New Freezeland Snowy Mammoth  · Frosty Retreat  · Hot Springs
Dimension/Dimension X Gladiator Attack
Recent Past/Medieval Mayhem Sushi Temple  · Wabi Sabi Wall  · Crumbling Castle
Present-Day/Futurama City Park  · Specter's Factory  · TV Tower
Present-Day 2/Specter Land Monkey Madness
Dimension/Dimension X Peak Point Matrix